A cast of leading animal biologists reflect on the enduring significance of Nobel laureate Niko Tinbergen's groundbreaking proposals for behavioural biology.An international cast of leading animal biologists reflect on the enduring significance of Tinbergen's groundbreaking proposals for modern behavioural biology. Includes a reprint of Nobel laureate Niko Tinbergen's original article on the famous 'four whys', after which each of the four questions are discussed in the light of contemporary evidence.An international cast of leading animal biologists reflect on the enduring significance of Tinbergen's groundbreaking proposals for modern behavioural biology. Includes a reprint of Nobel laureate Niko Tinbergen's original article on the famous 'four whys', after which each of the four questions are discussed in the light of contemporary evidence.Nobel laureate Niko Tinbergen laid the foundations for the scientific study of animal behavior with his work on causation, development, function and evolution. In this book, an international cast of leading animal biologists reflect on the enduring significance of Tinbergens groundbreaking proposals for modern behavioral biology. It includes a reprint of Tinbergens original article on the famous four whys and a contemporary introduction, after which each of the four questions are discussed in the light of contemporary evidence. Also discussed is the wider significance of recent trends in evolutionary psychology and neuroecology to integrate the four whys . With a foreword by one of Tinbergens most prominent pupils, Aubrey Manning, this wide-ranging book demonstrates that Tinbergens views on animal behavior are crucial for modern behavioral biology. It will appeal to graduate students and researchers in animal behavior, behavioral ecology and evolutionary biology.1. On aims and methods of ethology N Tinbergen; 2. Tinbergen's four questions and contemporary behavioural biology Jerry A. Hogan and Johan J. Bolhuis; 3. Causatl#F